Tumors as Evolutionary Engines: The Surprising Theory of Carcino-Evo-Devo

For decades, cancer has been viewed as a biological enemy. What if we've been wrong all along?

Evolutionary Biology Cancer Research Developmental Biology

Introduction: Rethinking Cancer's Role in Nature

For centuries, tumors have been viewed as unqualified biological villains—rogue cellular processes that destroy lives without purpose. But what if certain tumors actually played a constructive role in shaping the diversity of life? This revolutionary perspective comes from carcino-evo-devo, a theory proposing that hereditary tumors have served as evolutionary innovation engines throughout the history of multicellular life.

Developed primarily by Andrei P. Kozlov and his colleagues, this theory suggests that benign, hereditary tumors provided the raw cellular materials necessary for evolutionary breakthroughs. They may have offered extra cell masses where evolutionarily novel genes could be expressed, potentially leading to the emergence of new cell types, tissues, and organs 3 .

This theory connects three fundamental biological processes: cancer (neoplastic development), evolution (evo), and individual development (devo). By examining their interconnection, carcino-evo-devo aims to explain previously puzzling biological phenomena and potentially become a unifying biological framework 3 6 .

Cancer Biology
Carcino-Evo-Devo
Evolutionary Theory
Developmental Biology

What Is Carcino-Evo-Devo? Understanding the Core Theory

The carcino-evo-devo theory challenges conventional wisdom by proposing that hereditary tumors have contributed to progressive evolution. The central hypothesis, termed "evolution by tumor neofunctionalization," posits that tumors provided developing organisms with additional cellular real estate where evolutionary innovations could emerge 3 .

Key Principles
  • Tumor processes participate in the evolution of developmental pathways
  • Hereditary tumors provide extra cell masses for novel gene expression
  • Populations of tumor-bearing organisms served as transitional forms
  • Tumors may be considered search engines for new gene combinations
Historical Foundations
  • Carcinoembryonic antigens linking tumors and embryonic development
  • Evolutionary developmental biology (evo-devo) established field
  • "Embryonal rest" theory of cancer origins
  • "Hopeful monsters" concept of major evolutionary mutations

This perspective doesn't suggest that malignant, lethal cancers provide evolutionary advantages. Rather, it focuses on benign tumors or those at early developmental stages that don't kill their hosts but provide cellular material where evolutionary experimentation can occur 3 .

The Evolutionary Puzzle: Where Do New Biological Features Come From?

A fundamental question in evolutionary biology has long been: How do genetic innovations find expression in physical form during evolution? Traditional evolutionary theory explains how existing traits are selected, but has struggled to fully account for the origin of completely novel structures like the brain, liver, or complex sensory organs.

Carcino-evo-devo proposes a solution: tumors may provide the necessary cellular playgrounds where new genes can be expressed without disrupting existing functions. These additional cell masses serve as evolutionary test beds where nature can experiment with new genetic combinations 3 .

The theory helps explain the mechanism behind what the evolutionary biologist Richard Goldschmidt termed "hopeful monsters"—organisms with radical mutations that might occasionally prove advantageous. Rather than waiting for gradual, step-by-step mutations, significant evolutionary advances might occur through tumor-mediated expressions of novel genetic combinations 3 .
Evolutionary Mechanisms
  • Gradual Mutations Traditional
  • Gene Duplication Established
  • Tumor Neofunctionalization Carcino-Evo-Devo

The Science in Action: Testing the Theory Through TSEEN Genes

A compelling theory must generate testable predictions, and carcino-evo-devo has done exactly this. One of its most significant predictions involves what Kozlov's team terms TSEEN genes—Tumor-Specific, Evolutionarily-New genes 1 .

The Hypothesis

If tumors indeed provide cellular material for expressing evolutionarily novel genes, then we should find genes that are:

  • Evolutionarily recent in origin
  • Primarily expressed in tumor tissues
  • Largely absent from normal adult tissues
The Experimental Approach

Kozlov's laboratory embarked on a series of investigations to identify and characterize such genes 1 3 6 :

  • Computational analysis of human genome databases
  • Comparative genomics to determine evolutionary novelty
  • Experimental measurement of gene expression
  • Creation of specialized databases to catalog TSEEN genes

Key Findings: The PBOV1 Gene Case Study

One compelling example is the PBOV1 gene, discovered to be a human-specific gene with virtually no expression in normal adult tissues but significant expression in various cancers, including breast, prostate, and ovarian cancers 6 .

Table 1: Experimental Results for PBOV1 Gene Expression
Tissue Type Gene Expression Level Significance
Normal breast tissue Undetectable Baseline control
Breast cancer High Confirms tumor-specific expression
Normal prostate tissue Undetectable Baseline control
Prostate cancer Moderate to High Supports tumor-specificity pattern
Various other normal adult tissues Undetectable to Minimal Reinforces evolutionary novelty hypothesis
Table 2: Additional TSEEN Genes Confirming the Theory
Gene Name Evolutionary Status Expression Pattern Implications
BRACHYURY (T) Evolutionarily conserved High in tumors, low in normal tissues Ancient genes can show TSEEN pattern
HS.633957 cluster Evolutionarily new Predominantly in tumors Supports the core prediction
Multiple other TSEEN genes Evolutionarily new Various human cancers Pattern holds across gene families

The Researcher's Toolkit: Key Materials and Methods

To conduct this pioneering research, scientists required specialized biological materials and methodological approaches:

Table 3: Essential Research Tools in Carcino-Evo-Devo Studies
Research Tool Primary Function Examples from Studies
Tumor Biobanks Provide diverse tissue samples for comparison Collections of various human cancer and normal tissues
Genomic Databases Identify evolutionarily novel genes Human genome databases, comparative genomic tools
Gene Expression Assays Measure where and when genes are active PCR, microarrays, RNA sequencing
Animal Models Study tumor processes in evolutionary context Goldfish with hyperplastic head growths
Bioinformatics Tools Analyze evolutionary relationships between genes Phylogenetic analysis software, sequence alignment programs
Data Analysis

Computational approaches to identify evolutionary patterns in tumor genomics

Molecular Biology

Techniques to measure gene expression across different tissue types

Evolutionary Modeling

Frameworks to understand how tumors might contribute to evolutionary change

Beyond the Lab: Implications and Connections

The carcino-evo-devo theory reaches beyond basic biological research, offering fresh perspectives on several scientific fronts:

Relationship to Other Evolutionary Theories

This theory doesn't seek to replace established evolutionary models but rather to complement them. It offers explanations for phenomena that other theories struggle to fully account for, particularly the rapid emergence of evolutionary novelties 1 6 .

Challenges to Conventional Cancer Research

The theory emerges alongside other critical reevaluations of cancer biology. Recent technological advances, particularly in genome sequencing, have revealed paradoxes challenging the orthodox "cancer as a genetic disease" paradigm .

As Huang, Soto, and Sonnenschein note in their 2025 PLOS Biology essay, large-scale cancer genome sequencing has uncovered "immense diversity" in genetic mutations between patients with the same nominal tumor type, questioning a strictly deterministic view of cancer causation through specific oncogenic pathways .

Medical Implications

While primarily an evolutionary theory, carcino-evo-devo may eventually influence medical approaches. The discovery of TSEEN genes opens possibilities for new therapeutic targets and diagnostic approaches 1 3 . Understanding the deep evolutionary connections between development and cancer may provide fresh perspectives on cancer prevention and treatment.

Conclusion: A New Perspective on Life's Complexity

The carcino-evo-devo theory represents a paradigm shift in how we view both cancer and evolutionary processes. By suggesting that tumors have played a role in life's creative diversification, it challenges deeply entrenched views of cancer as an unqualified biological evil.

Though the theory continues to develop and requires further validation, it has already generated confirmed predictions and provided plausible explanations for previously mysterious biological phenomena. It reminds us that in science, seemingly settled questions—like the fundamental nature of cancer—can contain hidden depths waiting to be explored.

As Academician E.D. Sverdlov noted in his review of Kozlov's work, this theory "may become a unifying biological theory" 2 . By connecting individual development, evolutionary history, and neoplastic processes, it offers a more integrated vision of life's complexity—one where even disease processes may contain echoes of creative possibilities.

For further exploration: The four-part series "Theory of the Evolutionary Role of Hereditary Tumors (Carcino-Evo-Devo): History of Development and Current State" was published in Biology Bulletin Reviews throughout 2024.

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